姓 名：赵曦阳
性 别：男
出生年月：1981.03
职 称：教授/博士生导师
专 业：林学
学 科：林木遗传育种
手 机：15246668860
邮箱：zhaoxyphd@163.com
学习、工作经历
2022.07-至今 吉林农业大学，林学与草学学院，院长/教授/博士生导师
2021.10-2022.07 吉林农业大学，林学与草学学院，教授/博士生导师
2019.09-2021.10 东北林业大学，林学院，副主任/副教授/博士生导师
2018.06-2019.09 东北林业大学，林学院，国家重点实验室副主任/副教授
2015.09-2018.06 东北林业大学，林学院，副教授
2012.07-2015.09 东北林业大学，林学院，讲师
2010.07-2012.07 北京林业大学，生物学院，博士后
2005.09-2010.07 北京林业大学，林木遗传育种，博士
2000.09-2004.07 北京林业大学，生物科学，学士
荣誉称号
1. 吉林省“长白英才计划”科技创新领军人才，2024（吉林省最高人才）
2. 入选全球前2%顶尖科学家榜单，2023
3. 第十七届吉林省有突出贡献专家，2023
4. 吉林省C类人才，2023
5. 第十五届林业青年科技奖，2019
学术兼职
1. Journal of Forestry Research，编委，2020-至今
2. Forestry Research，副主编，2022-2024
3. 林业科学，编委，2024-2029
4. 吉林农业大学学报，编委，2024-2029
5. 植物研究，编委，2024-2029
社会兼职
1. 中国林学会青年工作委员会，副理事长，2024.06-至今
2. 中国林学会林木遗传育种分会，副秘书长，2018.06-至今
3. 吉林省林学会，副理事长，2023.05-至今
4. 吉林省青年科技工作者协会农业科学技术专业委员会，主任，2024.06-至今
5. 吉林省林草遗传育种重点实验室，主任，2022.11-至今
研究方向
围绕优质、高产与抗逆林木良种选育为目标，重点开展东北地区杨树、红松种质资源收集、创制与良种选育工作，主要包括以下3个研究方向：
（1）红松、杨树、胡桃楸等乡土树种种质资源收集与评价；
（2）红松、杨树、胡桃楸良种选育与高效培育；
（3）基于全基因组的红松、杨树、胡桃楸重要性状分子机理解析；
科研项目
1. 主持国家“十四五”重点研发课题“青杨类大径级工业资源材高效培育技术研究”（2021YFD2201204）；2021.12-2024.11；800万元
2. 主持农业生物育种2030国家科技重大专项“东北地区抗虫高产杨树新品种设计与培育”（2022ZD0401504）；2022.12-2025.12；360万元
3. 主持国家自然科学基金面上项目“白桦开花抑制因子BpFLC的调控机理研究”（31770712）；2018-2021；72万元
4. 主持国家自然科学基金青年项目“基于三倍体杨树染色体重排的育种体系探讨”（31500549）；2016.01-2018.12；26万元
5. 主持中央财政林业科技推广示范资金项目“吉林省中西部利用樟子松嫁接红松营建红松果林技术研究推广示范”（JLT2023-17）；2023.01-2025.12；50万元
6. 主持黑龙江省应用技术研究与开发计划项目“槭树类彩叶乡土树种良种选育与高效繁育技术研究”（GA20B402）；2020.08-2023.07；100万元
7. 主持中央高校基本科研业务费专项资金项目“东北地区红松和彩叶树种种质创新与推广示范”（2572020DR01）；2020-2023；100万元
8. 主持国家林业和草原局林业科技发展项目“红松种质资源遗传多样性评价”（KJZXSA2019046）；2019-2020；20万元
9. 主持“十三五”国家重点研发专项任务“吉林西部青杨类工业资源材高效培育技术研究”（2016YFD060040406）；2016-2020；82.11万元
10. 主持“十二五”国家科技计划课题子课题“白桦优树选择及种子园营建技术”（2012BAD21B0201-05）；2012-2016；35万元
11. 主持“十二五”国家科技计划课题子课题“落叶松木材相关候选基因SNP的开发”（2013AA102704-04）；2013-2017；24万元
12. 主持中央高校基本科研业务费专项资金项目“长白落叶松种子园优良亲本及子代评价选择”（2572017DA02）；2017-2019；35万元
发表文章
1. Pei Lei, Yaxuan Jiang, Yong Zhao, Mingquan Jiang, Ximei Ji, Le Ma, Guangze Jin, Jianxin Li, Subin Zhang, Dexin Kong, Xiyang Zhao*, Fanjuan Meng*. Functions of Basic Helix–Loop–Helix (bHLH) Proteins in the Regulation of Plant Responses to Cold, Drought, Salt, and Iron Deficiency: A Comprehensive Review. Journal of Agricultural and Food Chemistry. 2024, doi:10.1021/acs.jafc.3c09665.（IF=5.7; 1区TOP）
2. Yutong Ding, Jiajia Shen, Hanxi Li, Yue Sun, Tingbo Jiang, Xiangbo Kong, Rui Han, Chunli Zhao, Xinxin Zhang*, Xiyang Zhao*. Physiological and molecular mechanism of Populus pseudo-cathayana × Populus deltoides response to Hyphantria cunea. Pesticide Biochemistry and Physiology, 2024, doi:10.1016/ j.pestbp. 2024.105969. （IF=4.2; 1区TOP）
3. Yan Li, Runxian Song, Kewei Cai, Zhongyi Pang, Chengyuan Qian, Shujun Xu, Yu Zhang, Hui Bai, Wang Zhan, Ruijiang Xiao, Xiaona Pei, Rui Han*, Xiyang Zhao*. Exploring the molecular mechanisms of melatonin-induced tolerance to salt-alkali stress in Populus cathayana × canadansis 'Xinlin1′. Industrial Crops and Products, 2024, doi:10.1016/j.indcrop.2024.118638.（IF=5.6; 1区TOP）
4. Yan Li, Qiang Fu, Xin Li, Qinhui Zhang, Qiushuang Zhao, Yutong Ding, Jiajia Shen, Rui Han, Xiaona Pei, Xiyang Zhao*. Exploring the guardian of abiotic stress: Genome-wide identification of the basic helix-loop-helix transcription factor family in Juglans mandshurica. Scientia Horticulturae, 2024, doi:10.1016/ j.scienta. 2024.113154.（IF=5.6; 1区TOP）
5. Kewei Cai, Xiaoyu Xie, Lu Han, Junbo Chen, Jinwang Zhang, Hongtao Yuan, Jiajia Shen, Yishuang Ren, Xiyang Zhao*. Identification and functional analysis of the DOF gene family in Populus simonii: implications for development and stress response. Frontiers in Plant Science, 2024, doi:10.3389/fpls.2024.1412175. （IF=4.1; 1区TOP）
6. Yimin You, Liran Wang, Xiaoting Liu, Xuelai Wang, Luping Jiang, Changjun Ding, Weina Wang, Dawei Zhang, XiyangZhao*. Interspecific plant interaction structures the microbiomes of poplar-soil interface to alter nutrient cycling and utilization. Microbiology Spectrum, 2024, doi:10.1128/spectrum.03368-23. （IF=3.7; 1区TOP）
7. Jianxin Li, Subin Zhang, Pei Lei, Liyong Guo, Xiyang Zhao*, Fanjuan Meng*. Physiological and Proteomic Responses of the Tetraploid Robinia pseudoacacia L. to High CO2 Levels. International Journal of Molecular Sciences, 2024, doi:10.3390/ijms25105262.（IF=4.9; 3区）
8. Hongxing Wang, Luping Jiang, Feifan Zhang, Xiyang Zhao*. Effects of planting density and nitrogen fertilization on growth traits, leaf and wood characteristics of three poplar clones. Sustainability, 2024.（IF=3.3; 2区）
9. Hanxi Li, Jiajia Shen, Yutong Ding, Yuxi Li, Jiayu Du, Tingbo Jiang, Xiangbo Kong, Rui Han, Xixin Zhang*, Xiyang Zhao*. Transcriptomic and metabolomic analysis of poplar response to feeding by Hyphantria cunea. BMC plant biology, 2024, doi:10.1186/s12870-024-05631-2.（IF=4.3; 2区 TOP）
10. Ying Cui, Jiawei Yan, Luping Jiang, Junhui Wang, Manman Huang, Xiyang Zhao*, Shengqing Shi*. Needle and branch trait variation analysis and associated SNP loci mining in Larix olgensis. International Journal of Molecular Sciences, 2024, doi:10.3390/ijms251810212.（IF=4.9; 3区）
11. Kewei Cai, Dandan Zhang, Xiang Li, Qinhui Zhang, Luping Jiang, Yuxi Li, Runxian Song, Siqi Sun, Ruixue Guo, Rui Han, Xiaoqing Hu, Xudan Zhou*, Xiaona Pei*，Xiyang Zhao*. Exogenous phytohormone application and transcriptome analysis provides insights for adventitious root formation in Taxus cuspidata S. et Z. Plant Growth Regulation, 2023, doi:10.1007/s10725-022-00934-6.（IF=3.5; 3区）
12. Kewei Cai, Qiushuang Zhao, Hanxi Li, Qinhui Zhang, Yan Li, Rui Han, Tingbo Jiang, Xiaona Pei, Lina Zhang*, Xiyang Zhao*. Deciphering aroma formation during flowering in nectar tree (Tilia amurensis): Insights from integrated metabolome and transcriptome analysis. Forestry Research, 2023, doi:10.48130/FR-2023-0024.（2区）
13. Kewei Cai, Qiushuang Zhao, Jinwang Zhang, Hongtao Yuan, Hanxi Li, Lu Han, Xuebo Li, Kailong Li, Tingbo Jiang, and Xiyang Zhao*. Unraveling the guardians of growth: A comprehensive analysis of the aux/IAA and ARF gene families in Populus simonii. Plants, 2023, doi:10.3390/plants12203566. （IF=4; 2区）
14. David Kombi Kaviriri, Ling Yang, Chunli Zhao, Xiaona Pei, Zuoyi Fan, Jingyuan Wang, Liang Xu, Mulualem Tigabu, Xiangling You, Yuhua Li, Huan-Zhen Liu* and Xiyang Zhao*. Morphological growth performance and genetic parameters on Korean pine in Northeastern China. Silvae Genetica, 2023, doi:10.2478/sg-2023-0004.（IF=1.1; 4区）
15. Yan Li, Yujin Xu, Rui Han, Lin Liu, Xiaona Pei*, Xiyang Zhao*. Widely targeted metabolomic profiling combined with transcriptome analysis provides new insights into lipid biosynthesis in seed kernels of Pinus koraiensis. International Journal of Molecular Sciences, 2023, doi:10.3390/ijms241612887.（IF=4.9; 3区）
16. Yan Li, Minghui Zhao, Kewei Cai, Lin Liu, Rui Han, Xiaona Pei, Lina Zhang*, Xiyang Zhao*. Phytohormone biosynthesis and transcriptional analyses provide insight into the main growth stage of male and female cones Pinus koraiensis. Frontiers in Plant Science, 2023, doi:10.3389/fpls.2023.1273409.（IF=4.1; 1区TOP）
17. Qiushuang Zhao, Rui Han, Kewei Cai, Huiling Yan, Yan Li, Guanzheng Qu, Lin Liu*, Xiyang Zhao*. Identification and analysis of the CBF gene family in three species of Acer under cold stress. International Journal of Molecular Sciences, 2023, doi:10.3390/ijms24032088.（IF=4.9; 3区）
18. Minghui Zhao, Yan Li, Xinxin Zhang, Xiangling You, Haiyang Yu, Ruixue Guo*, Xiyang Zhao*. Genome-wide identification of AP2/ERF superfamily genes in Juglans mandshurica and expression analysis under cold stress. International Journal of Molecular Sciences, 2022, doi:10.3390/ijms232315225.（IF=4.9; 3区）
19. Xinxin Zhang, Yuxi Li, Huiling Yan, Kewei Cai, Hanxi Li, Zhiwei Wu, Jianguo Wu, Xiangdong Yang, Haichen Jiang, Qingcheng Wang, Guanzheng Qu, Xiyang Zhao*. Integrated metabolomic and transcriptomic analyses reveal different metabolite biosynthesis profiles of Juglans mandshurica in shade. Frontiers in Plant Science, 2022, doi:10.3389/fpls.2022.991874.（IF=4.1; 1区TOP）
20. Qinhui Zhang, Xinxin Zhang, Yuchun Yang, Lianfeng Xu, Jian Feng, Jingyuan Wang, Yongsheng Tang, Xiaona Pei*, Xiyang Zhao*. Genetic diversity of Juglans mandshurica populations in Northeast China based on SSR markers. Frontiers in Plant Science, 2022, doi:10.3389/fpls.2022.931578.（IF=4.1; 1区TOP）
21. Qinhui Zhang, Shihe Yu, Xiaona Pei, Qianchun Wang, Aijun Lu, Ying Cao, Mulualem Tigabu, Jian Feng, Xiyang Zhao*, Within-and between-population variations in seed and seedling traits of Juglans mandshurica, Journal of Forestry Research, 2022, doi:10.1007/s11676-021-01387-9.（IF=3.4; 2区）
22. Huiling Yan, Xinxin Zhang, Xiang Li, Xuelai Wang, Hanxi Li, Qiushuang Zhao, Peng Yin, Ruixue Guo, Xiaona Pei, Xiaoqing Hu, Rui Han*, Xiyang Zhao*. Integrated transcriptome and metabolome analyses reveal the anthocyanin biosynthesis pathway in AmRosea1 overexpression 84K poplar. Frontiers in Bioengineering and Biotechnology, 2022, doi:10.3389/fbioe.2022.911701. （IF=4.3; 3区）
23. Jiatong Wei, Xiaona Pei, Xiaoqing Hu, Siqi Sun, Chunli Zhao, Rui Han*, Xiyang Zhao*. Applications of transcriptome in conifer species. Plant Cell, Tissue and Organ Culture, 2022, doi:10.1007/s11240-022-02322-4.（IF=2.3; 3区）
24. Luping Jiang, Xiangzhu Xu, Qun Cai, Rui Han, Mulualem Tigabu, Tingbo Jiang*, Xiyang Zhao*. Variations in growth and photosynthetic traits of polyploid poplar hybrids and clones in Northeast China. Genes, 2022, doi:10.3390/genes13112161. （IF=2.8; 3区）
25. Hanxi Li, Yuxi Li, Xinxin Zhang, Kewei Cai, Yan Li, Qingcheng Wang, Guanzheng Qu, Rui Han*, Xiyang Zhao*. Genome-wide identification and expression analysis of the MADS-box gene family during female and male flower development in Juglans mandshurica. Frontiers in Plant Science, 2022, doi:10.3389/fpls.2022.1020706.（IF=4.1; 1区TOP）
26. Xiang Li, Kewei Cai, Zuoyi Fan, Jingyuan Wang, Lianfu Wang, Qi Wang, Lixing Wang, Xiaona Pei*, Xiyang Zhao*. Dissection of Transcriptome and Metabolome Insights into the Isoquinoline Alkaloid Biosynthesis during stem development in Phellodendron amurense (Rupr.). Plant Science, 2022, doi:10.1016/ j.plantsci. 2022.111461.（IF=4.2; 2区）
27. Xiang Li, Kewei Cai, Qinhui Zhang, Xiaona Pei, Song Chen, Luping Jiang, Zhiming Han, Minghui Zhao, Yan Li, Xinxin Zhang, Yuxi Li, Shikai Zhang, Su Chen, Guanzheng Qu, Mulualem Tigabu, Vincent L Chiang, Ronald Sederoff, Xiyang Zhao*. The manchurian walnut genome: insights into juglone and lipid biosynthesis. Gigascience, 2022, doi:10.1093/gigascience/giac057. （IF=11.8; 2区）
28. Yuxi Li, Xinxin Zhang, Yan Zhu, Kewei Cai, Hanxi Li, Qiushuang Zhao, Luping Jiang, Yan Li, Tingbo Jiang, Xiyang Zhao*. Physiological and transcriptomic analysis revealed the molecular mechanism of Pinus koraiensis responses to light. International Journal of Molecular Sciences, 2022, doi:10.3390/ijms232113608. （IF=4.9; 3区）
29. Xianbo Lu, Changjun Ding, Luping Jiang, Haiyang Yu, Rui Han, Jingwen Xu, Bin Li, Zhaoxiang Zheng, Chunming Li, Guanzheng Qu, Xiaona Pei*, Xiyang Zhao*. Evaluation of comprehensive effect of different agroforestry intercropping modes on poplar. Forests, 2022, doi:10.3390/f13111782.（IF=2.4; 2区）
30. Runxian Song, Faujiah Nurhasanah Ritonga, Haiyang Yu, Changjun Ding*, Xiyang Zhao*. Plant melatonin: Regulatory and protective role. Horticulturae, 2022, doi:10.3390/horticulturae8090810.（IF=3.1; 2区）
31. Anran Sun, Xiaona Pei, Shikai Zhang, Zhiming Han, Ying Xie, Guanzheng Qu, Xiaoqing Hu*, Mulualem Tigabu*, Xiyang Zhao*. Metabolome and transcriptome analyses unravels molecular mechanisms of leaf color variation by anthocyanidin biosynthesis in Acer triflorum. Horticulturae, 2022, doi:10.3390/horticulturae8070635.（IF=3.1; 2区）
32. Xiaoting Liu, Zhimin Lu, Jingui Wu, Yuchun Yang, Shuchun Li, Lin Wu, Qingshan Xu, Qi Yu, Li Zhao, Jian Sun, Wei Zheng, Mulualem Tigabu, Xiyang Zhao*. Comparison of genetic impact on growth and wood traits between seedlings and clones from the same plus trees of Pinus koraiensis. Journal of Forestry Research, 2022, doi:10.1007/s11676-022-01480-7.（IF=3.4; 2区）
33. Xiang Li, Rui Han, Kewei Cai, Ruixue Guo, Xiaona Pei, Xiyang Zhao*. Characterization of phytohormones and transcriptomic profiling of the female and male in florescence development in manchurian walnut (Juglans mandshurica Maxim.). International Journal of Molecular Sciences, 2022, doi:10.3390/ ijms23105433.（IF=4.9; 3区）
34. Zhiming Han, Xiangzhu Xu, Shikai Zhang, Qiushuang Zhao, Hanxi Li, Ying Cui, Xiao Li, Liran Wang, Su Chen, Xiyang Zhao*. Transcriptomics Profiling of Acer pseudosieboldianum Molecular Mechanism against Freezing Stress, International Journal of Molecular Sciences, 2022, doi:10.3390/ijms232314676.（IF=4.9; 3区）
35. Yuxi Li, Xinxin Zhang, Qinhui Zhang, Luping Jiang, Rui Han, Siqi Sun, Xiaoqing Hu, Xiaona Pei, Chunli Zhao, Xiyang Zhao*. G × E analysis of growth traits of Betula platyphylla clones at three separated sites in northeastern China. Phyton-International Journal of Experimental Botany, 2022, doi: 10.32604/ phyton. 2022.020518.（IF=1.3; 4区）
36. Heng Zhang, Shikai Zhang, Wenqi Song, Mulualem Tigabu, Meng Fu, Huifen Xue, Anran Sun, Minghui Zhao, Kewei Cai, Yan Li, Dean Xia, Chuanping Yang*, Xiyang Zhao*. Climate response of radial growth and early selection of Larix olgensis at four trials in northeast China. Dendrochronologia, 2022, doi:10.1016/j.dendro.2022.125955.（IF=2.7; 3区）
37. Xiang Li, Kewei Cai, Zhiming Han, Shikai Zhang, Anran Sun, Ying Xie, Rui Han, Ruixue Guo, Mulualem Tigabu, Ronald Sederoff, Xiaona Pei*, Chunli Zhao*, Xiyang Zhao*. Chromosome-level genome assembly for Acer pseudosieboldianum and highlights to mechanisms for leaf color and shape change. Frontiers in Plant Science, 2021, doi:10.3389/fpls.2022.850054. IF=4.1; 1区TOP）
38. Qinhui Zhang, Xiaona Pei, Lianfeng Xu, Xianbo Lu, Baoyang Wen, Yanlong Li, Liankui Wang, Guangzhi Dong, Wanling Shi, Xiaoqing Hu*, Xiyang Zhao*. Genetic improvement of Betula platyphylla Suk. in China: a review. Phyton-International Journal of Experimental Botany, 2021, doi: 10.32604/ phyton. 2022. 020372.（IF=1.3; 4区）
39. Qinhui Zhang, Xiaona Pei, Xianbo Lu, Chunli Zhao, Guangzhi Dong, Wanling Shi, Liankui Wang, Yanlong Li, Xiyang Zhao*, Mulualem Tigabu. Variations in growth traits and wood physicochemical properties among Pinus koraiensis families in Northeast China. Journal of Forestry Research, 2021, doi:10.1007/ s11676-022-01455-8.（IF=3.4; 2区）
40. Heng Zhang, Shikai Zhang, Shuopeng Chen, Dean Xia, Chuanping Yang*, Xiyang Zhao*. Genetic variation and superior provenances selection for wood properties of Larix olgensisat four trials. Journal of Forestry Research, 2021, doi:10.1007/s11676-021-01449-y.（IF=3.4; 2区）
41. Xiang Li, Kewei Cai, Qiushuang Zhao, Hanxi Li, Xuelai Wang, Mulualem Tigabu, Ronald Sederoff, Wenjun Ma*, Xiyang Zhao*. Morphological and comparative transcriptome analysis of three species of five-needlepines: insights into phenotypic evolution and phylogeny. Frontiers in Plant Science, 2021, doi:10.3389/fpls.2022.795631.（IF=4.1; 1区TOP）
42. Kewei Cai, Xueyan Zhou, Xiang Li, Ye Kang, Xiaoming Yang, Yonghong Cui, Guangyan Li, Xiaona Pei*, Xiyang Zhao*. Insight into the multiple branches traits of a mutant in Larix olgensis by morphological, cytological, and transcription alanalyses. Frontiers in Plant Science, 2021, doi:10.3389/ fpls.2021. 787661.（IF=4.1; 1区TOP）
43. Shikai Zhang, Wang Zhan, Anran Sun, Ying Xie, Zhiming Han, Xibin Qu, JiayiWang, Laifu Zhang, MingshunTian, Xuhong Pang, Jinbao Zhang, Xiyang Zhao*. Combined transcriptome and metabolome integrated analysis of Acer mandshuricum to reveal candidate genes involved in anthocyanin accumulation. Scientific reports, 2021, doi:10.1038/s41598-021-02607-2.（IF=3.8; 2区）
44. Yan Li, Xiang Li, Jiatong Wei, Kewei Cai, Hongzhi Zhang, Lili Ge, Zengjun Ren, Chunli Zhao*, Xiyang Zhao*. Genome-wide identification and analysis of the WRKY gene family and cold stress response in Acer truncatum. Genes, 2021, doi:10.3390/genes12121867.（IF=2.8; 3区）
45. Jiatong Wei, Xiang Li, Huazhe Xu, Yuliang Wang, Chuanhai Zang, Jingwen Xu, Xiaona Pei*, Xiyang Zhao*. Evaluation of the genetic diversity of Pinus koraiensis by EST-SSR and its management, utilization and protection. Forest Ecology and Management, 2021, doi:10.1016/j.foreco.2021.119882.（IF=3.7; 1区TOP）
46. Fang Wang, Song Chen, Kewei Cai, Zhimin Lu, Yuchun Yang, Mulualem Tigabu, Xiyang Zhao*. Transcriptome sequencing and gene expression profiling of Pinus sibirica under different cold stresses. Breeding Science, 2021, doi:10.1270/ jsbbs.21009.（IF=2; 4区）
47. Shikai Zhang, Wang Zhan, Heng Zhang, Shuanglin Song, Xibin Qu, Jiayi Wang, Mulualem Tigabu, Xiang Li, Qinhui Zhang, Minghui Zhao, Huanzhen Liu*, Xiyang Zhao*. Geneticim provement in Juglans mandshurica and its uses in China: current status and future prospects. Phyton-international Journal of Experimental Botany, 2021, doi:10.32604/phyton.2022.018321.（IF=1.3; 4区）
48. Xiang Li, Kewei Cai, Xiaona Pei, Yan Li, Yanbo Hu, Fanjuan Meng, Xingshun Song, Mulualem Tigabu, Changjun Ding*, Xiyang Zhao*. Genome-wide identification of NAC transcription factor family in Juglans mandshurica and their expression analysis during the fruit development and ripening. International Journal of Molecular Sciences, 2021, doi:10.3390/ ijms 222212414.（IF=4.9; 3区）
49. Xiang Li, Yan Li, Minghui Zhao, Yanbo Hu, Fanjuan Meng, Xingshun Song, Mulualem Tigabu, Vincent L. Chiang, Ronald Sederoff, Wenjun Ma, Xiyang Zhao*. Molecular and metabolic insights into anthocyanin biosynthesis for leaf color change in chokecherry (Padus virginiana). International Journal of Molecular Sciences, 2021, doi:10.3390/ijms221910697.（IF=4.9; 3区）
50. Qinhui Zhang, Shihe Yu, Xiaona Pei, Qianchun Wang, Aijun Lu, Ying Cao, Mulualem Tigabu, Jian Feng, Xiyang Zhao*. Within‑and between‑population variations in seed and seedling traits of Juglans mandshurica. Journal of Forestry Research, 2021, doi:10.1007/s11676-021-01381-1.（IF=3.4; 2区）
51. Yan Li, Xiang Li, Minghui Zhao, Zhongyi Pang, Jiatong Wei, Mulualem Tigabu, Vincent L. Chiang, Heike Sederoff, Ronald Sederoff, Xiyang Zhao*. An overview of the practices and management methods for enhancing seed production in conifer plantations for commercial use. Horticulturae, 2021, doi:10.3390/ horticulturae 7080252.（IF=3.1; 2区）
52. Luping Jiang, Xiaona Pei, Yanbo Hu, Vincent L. Chiang, Xiyang Zhao*. Effects of environment and genotype on growth traits in poplar clones in Northeast China. Euphytica, 2021, doi:10.1007/s10681-021-02894-w.（IF=1.6; 3区）
53. Liping Xu, Yanbo Hu, Guangze Jin, Pei Lei, Liqun Sang, Qiuxiang Luo, Zhi Liu, Fachun Guan, Fanjuan Meng*, Xiyang Zhao*. Physiological and proteomic responses to drought in leaves of Amygdalus mira (Koehne) Yü et Lu. Frontiers in Plant Science, 2021, doi:10.3389/fpls.2021.620499.（IF=4.1; 1区TOP）
54. Lijiao Sun, Xiaoyu Zhao, Jing Ren, Shaopeng Yan, Xiyang Zhao*, Xingshun Song*. Overexpression of Cerasus humilis ChAOX2 improves the tolerance of Arabidopsis to salt stress. 3 Biotech, 2021, doi:10.1007/s13205- 021- 02871-z. （IF=2.6; 4区）
55. Shuchun Li, Heng Zhang, Zuoyi Fan, Deyang Liang, Haiqi Sun, Yue Li, Xiyang Zhao*. Genetic test and early selection in full-sib families of Pinus koraiensis. Scandinavian Journal of Forest Research, 2021, doi:10.1080/02827581. 2021. 1919752.（IF=3.4; 2区）
56. Xiang Li, Minghui Zhao, Yujin Xu, Yan Li, Mulualem Tigabu, Xiyang Zhao*. Genetic diversity and population differentiation of Pinus koraiensis in China. Horticulturae, 2021, doi:10.3390/horticulturae7050104.（IF=3.1; 2区）
57. David Kombi Kaviriri, Huanzhen Liu*, Xiyang Zhao*. Estimation of genetic parameters and wood yield selection index in aclonal trial of Korean Pine (Pinus koraiensis) in northeastern China. Sustainability, 2021, doi:10.3390/su13084167. （IF=3.3; 3区）
58. Huiling Yan, Xiaona Pei, Heng Zhang, Xiang Li, Xinxin Zhang, Minghui Zhao, Vincent L. Chiang, Ronald Ross Sederoff, Xiyang Zhao*. MYB-mediated regulation of anthocyanin biosynthesis. International Journal of Molecular Sciences, 2021, doi:10.3390/ijms 22063103.（IF=4.9; 3区）
59. Heng Zhang, Xueyan Zhou, Wei Gu, Lidong Wang, Wei Li, Yuanke Gao, Liguo Wu, Xiaowei Guo, Mulualem Tigabu, Dean Xia, Vincent L. Chiang, Chuanping Yang*, Xiyang Zhao*. Genetic stability of Larix olgensis provenances planted in different sites in northeast China. Forest Ecology and Management, 2021, doi:10.1016/j.foreco.2021.118988.（IF=3.7; 1区TOP）
60. Xiaona Pei, Luping Jiang, Ammar khalil Mohamed Ahmed, Hongying Yu, Rizheng Chong, Xiangling You, Xiyang Zhao*. Growth variations and stability analyses of seven poplar clones at three sites in Northeast China. Journla of Forestry Research, 2020, doi:10.1007/s 11676- 020-01210-x. （IF=3.4; 2区）
61. Yanbo Hu, Manzer H. Siddiqui, Chunming Li, Luping Jiang, Heng Zhang, Xiyang Zhao*. Polyamine metabolism photorespiration, and excitation energy allocation in photosystem П are potentially regulatory hubs in poplar adaptation to soil nitrogen availability. Frontiers in Plant Science, 2020，doi:10.3389/ fpls.2020.01271.（IF=4.1; 1区TOP）
62. Minghui Zhao, Xiang Li, Xinxin Zhang, Heng Zhang, Xiyang Zhao*. Mutation mechanism of leaf color in plants: a review. Forests, 2020, doi:10.3390/f11080851. （IF=2.4; 2区）
63. Yanbo Hu, Chunming Li, Luping Jiang, Deyang Liang, Xiyang Zhao*. Growth performance and nitrogen allocation within leaves of two poplar clones after exponential and conventional nitrogen applications. Plant Physiology and Biochemistry, 2020, doi:10.1016/j.plaphy.2020.06.053.（IF=6.1; 2区TOP）
64. Xiang Li, Xiaoting Liu, Jiatong Wei, Yan Li, Mulualem Tigabu, Xiyang Zhao*. Development and transferability of EST-SSR markers for Pinus koraiensis from cold-stressed transcriptome through illumina sequencing. Genes, 2020, doi:10.3390/genes11050500.（IF=2.8; 3区）
65. David Kombi Kaviriri, Qinhui Zhang, Xinxin Zhang, Luping Jiang, Jinpeng Zhang, Jingyuan Wang, Damase P. Khasa, Xiangling You, Xiyang Zhao*. Phenotypic variability and genetic diversity in a Pinus koraiensis clonal trial in Northeast China. Genes, 2020, doi:10.3390/genes11060673.（IF=2.8; 3区）
66. Heng Zhang, Yuhangyi Zhang, Dawei Zhang, Lihu Dong, Kejian Liu, Ying Wang, Chuanping Yang, Vincent L. Chiang, Mulualem Tigabu, Xiyang Zhao*. Progeny performance and selection of superior trees within families in Larix olgensis. Euphytica, 2020, doi:10.1007/s10681- 020-02596-9.（IF=1.6; 3区）
67. David Kombi Kaviriri, Xiaoting Liu, Zuoyi Fan, Jingyuan Wang, Qi Wang, Lianfu Wang, Lixing Wang, Damase Khasa, Xiyang Zhao*. Genetic variation in growth and cone traits of Pinus Koraiensis half-sib families in northeast China. Phyton-International Journal of Experimental Botany, 2020, doi:10.32604/ phyton. 2020.08409.（IF=1.3; 4区）
68. Xiang Li, Xiaoting Liu, Jiatong Wei, Yan Li, Mulualen Tigabu, Xiyang Zhao*. Genetic improvement of Pinus koraiensis in China: current situation and future prospects. Forests. 2020, doi:10.3390/f11020148.（IF=2.4; 2区）
69. Yanyan Pan, Luping Jiang, Guiyou Xu, Jiaqi Li, Biying Wang, Yuxi Li, Xiyang Zhao*. Evaluation and selection analyses of 60 Larix kaempferi clones in four provenances based on growth traits and wood properties.Tree Genetics&Genomes. 2020, doi:10.1007/s11295- 020-1420-z.（IF=1.9; 2区）
70. David Kombi Kaviriri, Yuxi Li, Dawei Zhang, Hongtao Li, Zuoyi Fan, Jingyuan Wang, Lianfu Wang, Qi Wang, Deqiu Wang, Vincent L. Chiang, Xiyang Zhao*. Clonal variations in cone, seed and nut traits in a Pinus koraiensis seed orchard in Northeast China. Journal of Forestry Research. 2020, doi:10.1007/s11676-019-01094-6.（IF=3.4; 2区）
71. Fang Wang; Song Chen, Deyang Liang, Guanzheng Qu, Su Chen, Xiyang Zhao*. Transcriptomic analyses of Pinus koraiensis under different cold stresses. BMC Genomics. 2020, doi:10.1186/s12864-019-6401-y.（IF=2.1; 4区）
72. Ammar Khalil Mohamed Ahmed, Luping Jiang, Fang Wang, Song Chen, Xueyan Zhou, Xiaona Pei, Xiyang Zhao*, Guanzheng Qu*. Variation analysis of growth traits of four poplar clones under different water and fertilizer management. Journal of Forestry Research, 2019, doi:10.1007/s11676- 019-00888-y.（IF=3.4; 2区）
73. Ammar Khalil Mohamed Ahmed, Zhixiyang Fu, Changjun Ding, Luping Jiang, Xidong Han, Aiguo Yang, Yinghui Ma, Xiyang Zhao*. Growth and wood properties of a 38-year-old Populus simonii × P. nigra plantation established with different densities in semi-arid areas of northeastern China. Journal of Forestry Research, 2019, doi:10.1007/s11676-019- 00887-z.（IF=3.4; 2区）
74. Deyang Liang, Biying Wang, Shuanglin Song, Jingyuan Wang, Lianfu Wang, Qi Wang, Xuanbai Ren, Xiyang Zhao*. Analysis of genetic effects on a complete diallel corss test of Pinus koraiensis. Euphytica, 2019, doi:10.1007/s10681-019-2414-5.（IF=1.6; 3区）
75. Song Chen, Xin Lin, Dawei Zhang, Qi Li, Xiyang Zhao*, Su Chen*. Genome-wide analysis of NAC gene family in Betula pendula. Forests, 2019, doi:10.3390/f10090741.（IF=2.4; 2区）
76. Yanyan Pan, Xiaona Pei, Fuwei Wang, Chenglu Wang, Lili Shao, Lihu Dong, Xiyang Zhao*, GuanzhengQu. Forward, backward selection and variation analysis of growth traits in half-sib Larix kaempferi families. Silvae Genetica, 2019, doi:10.2478/sg-2019-0001.（IF=1.1; 4区）
77. Zhixin Li, Xiaona Pei, Shaopeng Yin, Xiangbo Lang, Xiyang Zhao*, GuanZheng Qu. Plant hormone treatments to alleviate the effects of salt stress on germination of Betula platyphylla seeds. Journal of Forestry Research, 2018, doi:10.1007/ s11676-018-0661-2.（IF=3.4; 2区）
78. Fang Wang, Deyang Liang, Xiaona Pei, Qinhui Zhang, Peng Zhang, Jianqiu Zhang, Zhimin Lu, Yuchun Yang, Guifeng Liu, Xiyang Zhao*. Study on the physiological indices of Pinus sibirica and Pinus koraiensis seedlings under cold stress. Journal of Forestry Research, 2018, doi:10.1007/s11676-018-0833-0.（IF=3.4; 2区）
79. Fang Wang, Qinhui Zhang, Yonggang Tian, Shuchen Yang, Hongwu Wang, Liankui Wang, Yanlong Li, Peng Zhang, Xiyang Zhao*. Comprehensive assessment of growth traits and wood properties in half-sib Pinus koraiensis families. Euphytica, 2018, doi:10.1007/s10681-018-2290-4.（IF=1.6; 3区）
80. Chunming Li, Hui Xia, Hui Bai, Hongmei Wang, Yajuan Xing, Xiyang Zhao*, Xiaomei Sun. Genetic variation of height growth rhythm between clones of Larix kaempferi × L. gmelini based on logistic models. Journal of Forestry Research, 2017, doi:10.1007/s11676-017-0558-5.（IF=3.4; 2区）
81. Deyang Liang, Changjun Ding, Guanghao Zhao, Weiwei Leng, Min Zhang, Xiyang Zhao*, Guanzheng Qu. Variation and selection analysis of Pinus koraiensis clones in northeast China. Journal of Forestry Research, 2017, doi:10.1007/s 11676-017-0471-y.（IF=3.4; 2区）
82. Yanyan Pan, Shuchun Li, Chenglu Wang, Wenjun Ma, Guiyou Xu, Lili Shao, Kailong Li, Xiyang Zhao*, Tingbo Jiang*. Early evaluation of growth traits of Larix kaempferi clones. Journal of Forestry Research, 2017, doi:10.1007/s11676-017-0492-6.（IF=3.4; 2区）
83. Deyang Liang, Xinxin Zhang, Chao Wang, Xiuwei Wang, Kailong Li, Guifeng Liu, Xiyang Zhao*, Guan-Zheng Qu. Evaluation of Betula platyphylla families based on growth and wood property traits. Forest Science, doi:10.1093/forsci/fxy027. （IF=1.5; 4区）
84. Shaopeng Yin, Zhenhai Xiao, Guohui Zhao, Xin Zhao, Xiaoyang Sun, Ying Zhang, Fuwei Wang, Shuchun Li, Xiyang Zhao*, Guan-Zheng Qu. Variation analyses of growth and wood properties of Larix olgensis clones in China. Journal of Forestry Research, 2016, doi:10.1007/s11676-016-0359-2.（IF=3.4; 2区）
85. Xiyang Zhao, Tangchun Zheng, Longting Shao, Zhenhai Xiao, Fuwei Wang, Shuchun Li, Lina Zang, Mi Zheng, Ying Li*, Guanzheng Qu*. Variation analysis of physiological traits in Betula platyphylla overexpressing TaLEA-ThbZIP gene under salt stress. PLoS One, 2016, doi:10.1371/journal.pone.0164820.（IF=2.9; 3区）
86. Xiyang Zhao, Hui Xia, Xiuwei Wang, Chao Wang, Deyang Liang, Kailong Li*, Guifeng Liu*. Variance and stability analyses of growth characters in half-sib Betula platyphylla families at three different sites in China. Euphytica, 2016, doi:10.1007/s10681-015-1617-7.（IF=1.6; 3区）
87. Dongjing Si, Mi Zheng, Ying Li, Guanzheng Qu, Xiyang Zhao*. Proteins involved in salt response mechanisms in Poplar (PopulusSPP.). Current Proteomics, 2016, doi:10.2174/1570164613666161019143019.（IF=0.5; 4区）
88. Hui Xia, Guohui Zhao, Liansheng Zhang, Xiaoyang Sun, Shaopeng Yin, Deyang Liang, Ying Li, Mi Zheng, Xiyang Zhao*. Genetic and variation analyses of growth traits of half-sib Larix olgensis families in northeastern China. Euphytica, 2016, doi:10.1007/s10681-016-1765-4.（IF=1.6; 3区）
89. Mengran Liu, Shaopeng Yin, Dongjing Si, Longting Shao, Ying Li, Mi Zheng, Fuwei Wang, Shuchun Li, Guifeng Liu, Xiyang Zhao*. Variation and genetic stability analyses of transgenic TaLEA poplar clones from four different sites in China. Euphtyica, 2015, doi:10.1007/s10681-015-1471-7.（IF=1.6; 3区）
90. Xiyang Zhao, Ying Li, Mi Zheng, Xiuyan Bian, Mengran Liu, Yanshuang Sun, Jing Jiang, Fuwei Wang, Shuchun Li, Yonghong Cui, Guifeng Liu, Chuanping Yang*. Comparative analysis of growth and photosynthetic characteristics of different ploidy level (Populus simonii × P. nigra) × (P. nigra × P. simonii) Hybrid Clones. PLoS One, 2014, doi:10.1371/journal.pone.0119259. (IF=2.9; 3区）
91. Xiyang Zhao*, Huiquan Zheng, Shanwen Li, Chuanping Yang, Jing Jiang, Guifeng Liu. The rooting of poplar cuttings: a review. New Forests, 2014, doi:10.1007/ s11056-013-9389-1. (IF=1.9; 3区）
92. Xiyang Zhao, Cheng Wang, Shuchun Li, Wei Hou, Shiqing Zhang, Guijie Han, Deng Pan, Ping Wang, Yufei Cheng, Guifeng Liu*. Genetic variation and selection of introduced provenances of Siberian pine (Pinus sibirica) in frigid regions of the greater xing’an range, northeast China. Journal of Forestry Research, 2014, doi:10.1007/s11676-014-0494-6.（IF=3.4; 2区）
93. Xiyang Zhao, Xiuyan Bian, Zhixin Li, Xiuwei Wang, Chengjun Yang, Guifeng Liu, Jing Jiang, Kentbayev Y, Kentbayeva B, Chuanping Yang*. Genetic stability analysis of introduced Betula pendula, Betula kirghisorum, and Betula pubescens families in saline-alkali soil of northeastern China. Scandinavian Journal of Forest Research, 2014, doi:10.1080/ 02827581. 2014. 960892.（IF=1.8; 3区）
94. Xiyang Zhao, Xiuyan Bian, Mengran Liu, Zhixin Li, Ying Li, Mi Zheng, Wenhua Teng, Jing Jiang, Guifeng Liu*. Analysis of genetic effects on a complete diallel cross test of Betula Platyphylla. Euphytica, 2014, doi:10.1007/s10681-014 -1147-8.（IF=1.6; 3区）
95. Xiyang Zhao*, Wei Hou, Huiquan Zhen, Zhiyi Zhang. Analyses of genotypic variation in white poplar clones at four sites in china. Silvae Genetica, 2013, doi:10.1515/sg-2013-0023.（IF=1.1; 4区）
96. Xiyang Zhao, Chunfa Tong, Xiaoming Pang, Zhong Wang, Yunqian Guo, Fang Du, Rongling Wu*. Functional mapping of ontogenyin flowering plants. Briefings in Bioinformatics, 2011, doi:10.1093/bib/bbr054.（IF=6.8; 1区TOP）
成果与奖励
1. 吉林省红松良种选育与推广示范，梁希林业科技进步奖三等奖，2019（1）
2. 临江红松种子园优良亲本及子代的选择，吉林省科学技术三等奖，2018（3）
3. 山地杨树高效培育技术研究，黑龙江省科学技术三等奖，2015（6）
4. 红松种质资源库营建与良种选育技术，吉林省科技成果，2024（1）
5. 长白落叶松短枝簇生芽变枝变异机理研究，吉林省科技成果，2024（1）
6. 松辽平原杨树栽培区新林1号杨大径材高效培育技术研究，吉林省科技成果，2024（1）
7. 松辽平原杨树大径材培育全周期复合经营模式研究，吉林省科技成果，2024（1）
8. 长白落叶松良种选育与种子园升级换代技术，吉林省科技成果，2024（1）
9. 林木种子园子代亲本检测技术研究，吉林省科技成果，2024（1）
10. 东北地区青杨类杨树壮苗培育关键技术，中科评字，2021（1）
专著
1. 赵曦阳，邬荣领. 白杨杂交育种，中国林业出版社，2013
2. 赵曦阳，李志新. 长白落叶松遗传改良，中国林业出版社，2019
3. 王芳，赵曦阳. 低温胁迫下红松与西伯利亚红松生理及分子响应，2020
4. 裴晓娜，赵曦阳. 东北半干旱地区杨树高效栽培技术研究，2020
专利与软著
1. 赵曦阳、王宏星、裴晓娜、由义敏、蒋路平. “杨树大径材高效培育病虫害防 治系统V1.0”. 软件著作权，登记号：2024SR1430762, 2024-09
2. 赵曦阳、裴晓娜、韩锐、胡晓晴、孙思琦、赵春莉。“一种获取银中杨非整倍体植株的方法”. 专利号：ZL 2022 1 0174693.0，2023-04
3. 赵曦阳、裴晓娜、韩锐、刘琳、赵秋爽、蔡柯伟、姜廷波、孔祥波. “一种光肩星天牛诱控实验用的装置”. 专利号：ZL 2023 2 0207617.5，2023-07
4赵曦阳、裴晓娜、韩锐、刘琳、赵秋爽、蔡柯伟、姜廷波、孔祥波. “一种美国白蛾诱捕装置”. 专利号：ZL 2023 2 0207630.0，2023-07
5. 赵曦阳、裴晓娜、韩锐、刘琳、赵秋爽、蔡柯伟、姜廷波、孔祥波. “一种危害杨树的天牛捕捉器”. 专利号：ZL 2023 2 0207611.8，2023-07
6. 赵曦阳、陈嵩、李嘉琪、张恒、李峪曦、周雪燕、赵男、刘晓婷、胡柏策、张秦徽. “一种组合式树干涂白刷”. 专利号：ZL 2019 2 0390353.5，2019-11
7. 赵曦阳、张恒、赵男、陈嵩、胡柏策、周雪燕、李峪曦、李嘉琪、刘晓婷、张秦徽. “具有筛除落叶功能的林木种子采集装置”. 专利号：ZL 2018 2 2048391.0，2019-07
8. 赵曦阳、刘晓婷、胡柏策、周雪燕、赵男、李峪曦、陈嵩、李嘉琪、张恒、王壁莹. “一种林业用取果器”. 专利号：ZL 2018 2 2048393. X
9. 赵曦阳、周雪燕、李峪曦、刘晓婷、李嘉琪、赵男、胡柏策、张恒、陈嵩、王壁莹.“银杏树苗生长矫正装置”.专利号：ZL 2018 2 1972509.2
10. 赵曦阳、曲冠证、李莹、郑密、李开隆、邵龙婷. “一种高效便捷的温室扦插白桦生根方法”. 专利号：ZL 2016 1 0131265.4
11. 赵曦阳、裴晓娜、潘艳艳、王芳、李树春、孙晓阳、张连生、王连福、王琪、刘克俭. “一种杨树飞絮处理装置”. 专利号：ZL 2017 2 1036293.4
12. 赵曦阳、裴晓娜、张秦徽、王碧莹、李志新、李莹、郑密、李开隆、由香玲、曲冠证.“一种林业专用的便携式多自由度喷药装置”. 专利号：ZL 2017 2 1204047.5
13. 赵曦阳、李莹、裴晓娜、郑密、梁德洋、蒋路平、王芳、王璧莹、张秦徽、索慧英.“基于激光测距传感器的树木胸径测量装置”.专利号：ZL 2017 2 1923554.4
14. 赵曦阳、王福维、李树春、王井源、潘艳艳、冷伟伟、肖德胜、赵昕、曹磊.“一种手推式自动栽苗器”.专利号： ZL 2016 2 1247054.9
15. 赵曦阳、李树春、王福维、赵国辉、潘艳艳.“种子收集网片及利用种子收集网片形成的种子收集装置”.专利号：ZL 2016 2 034 0961.1
16. 赵曦阳、郑密、李志新、曲冠证、由香玲、梁德洋、蒋路平、张贵新、李开隆.“一种定量施肥的栽苗器”.专利号：ZL 2016 2 1254857.7
17. 赵曦阳、李莹、曲冠证、郑密、李春明、邵龙婷、李开隆.“一种打孔式林木施肥机”.专利号：ZL 2014 2 0033945.9
18. 赵曦阳、李莹、曲冠证、郑密、李春明、邵龙婷、李开隆.“一种林木测高仪”.专利号：ZL 2014 2 0020068.1
新品种与良种
1. 国审（认）良种：白桦草河口种源、白桦清源种源、白桦帽儿山种源、白桦小 北湖种源、白桦家系1-7、白桦家系3-12、白桦家系4-7、白桦家系4-13
2. 省审（认）良种：吉祥1号杨、龙井市红松PK06、PK27、PK29、PK34、PK20
3. 林木新品种：吉瑞1号
标准
1. 赵曦阳、蒋立峰、吴军、孟鑫、魏亚松、李艳萍、周志军、王洪梅、赵明辉、张金玲. “红松种质资源调查、保存及评价技术规程”，黑龙江省地方标准，标准号：DB23/T 3172－2022
2. 赵曦阳、吴军、娄宇、孟鑫、魏亚松、李艳萍、王洪梅、周志军、张世凯、紫瑜、韩笑. “白牛槭种质资源调查与评价技术规程”黑龙江省地方标准.
标准号：DB23/T 3171—2022
